Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group

The large, widespread, Southeast Asian genus Glyphiulus Gervais, 1847 is reviewed, with special reference to the granulatus-group, established for the species whose primary characteristics are that the males have sternum 1 devoid of median structures, but supplied with two widely separated prongs, at the base of which are often 1- or 2-segmented vestigial legs, along with the carinotaxy patterns of the collum and metaterga which are highly characteristic and often stable. This group currently encompasses 20 species, all keyed here, including the redescribed pantropical species G. granulatus (Gervais, 1847), as well as G. subgranulatus n. sp., G. paragranulatus n. sp., G. semigranulatus n. sp., G. beroni n. sp., G. deharvengi n. sp., G. pergranulatus n. sp., G. latellai n. sp. and G. basalis n. sp., all from caves in southern China, and G. bedosae n. sp. from a cave in Laos. The following two synonymies are established: G. granulatus (Gervais, 1847) = G. vulgatus Zhang & Li, 1982, and G. septentrionalis Murakami, 1975 = G. multicarinus Zhang & Li, 1982.