Santonian Angiosperm-dominated leaf-assemblage from Piolenc (Vaucluse, Sud-Est de la France)

Lower Santonian sandy to coaly-clayey beds, collected near the village of Piolenc (Vaucluse) contain amber bearing insects and extremely well-preserved leaf cuticle compressions of Corystospermales, Cycadales, Bennettitales, Gymnosperms and mostly Angiosperms. The sedimentological and taphonomic studies indicate that the clays bearing the fossil cuticles were deposited along the Alpine Sea shore into supratidal ponds or marshes around which the plants studied herein grew. Moreover, the 'nearest living relative' and 'foliar physiognomy' methods, and the sedimentological and palaeontological studies agree with the occurrence of a warm and humid, tropical climate, including a dry season.