Scorpions of the Réserve spéciale d'Ankarana, Madagascar, with particular reference to cave-dwelling animals and the description of two new species (Arachnida, Scorpiones)

A synopsis is presented of the scorpion fauna documented in the Réserve spéciale d’Ankarana in northern Madagascar, an area of limestone karst. Two new species of liochelid scorpions, Opisthacanthus milloti n. sp. and Opisthacanthus pauliani n. sp., are described based on specimens collected in the reserve. One of these, O. pauliani n. sp. may represent the second troglobitic scorpion species found in Madagascar, and the first one belonging to the genus Opisthacanthus Peters, 1961 anywhere in the world.