Inferences from molecular data (rps4 and trnL-F) and morphological features of the Chilean endemic Aptychella chilensis reveal that the species should be excluded from Aptychella and the Pylaisiadelphaceae, and transferred to Ombronesus in the Ptychomniaceae. The new combination Ombronesus chilensis (Herzog) Frank Müll. & H. Akiyama is proposed with the new synonym O. stuvensis N.E. Bell, N. Pedersen & A.E. Newton. A lectotype for A. chilensis is selected.