A new elasmotheriine (Mammalia, Rhinocerotidae) from the Early Miocene of Spain

A new species of Hispanotherium from the Early Miocene of Spain is named. Its phylogenetic relationships within Elasmotheriina are discussed owing to a cladistic analysis. H. grimmi Heissig, 1974 and H. beonense Antoine, 1997 are consequently integrated in the genus Hispanotherium , together with the type species H. matritense and the new species, which differs from other ones by several dental and postcranial features. The westward dispersal of the Elasmotheriina from Asia toward Western Europe during the Early Miocene is hypothesized.