A contribution to the study of hornworts and liverworts in Tunisia: a checklist and ecology of Kroumirian species

Studies of the Tunisian bryoflora are disparate and mostly quite outdated. In order to enhance and update our knowledge, a survey of Anthocerotophyta and Marchantiophyta of Kroumiria was carried out in May 1-5, 2017, recording two hornworts and 30 liverworts including Riccia gougetiana Durieu & Mont. var. armatissima Levier ex Müll.Frib., new to the area. The liverwort and hornwort flora of Kroumiria now stands at 80 reliably recorded taxa, of which 22 species may be threatened. Numerical analyses were used to specify the requirements of the 32 listed taxa for substrate, light, temperature and humidity and to distinguish seven different ecological groups ranging from the hydro-thermo-photophile to the mesohygro-sciaphile.